He is no fool who gives what he cannot keep to gain that which he cannot lose怎么理解

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He is no fool who gives what he cannot keep to gain that which he cannot lose怎么理解

He is no fool who gives what he cannot keep to gain that which he cannot lose怎么理解
He is no fool who gives what he cannot keep to gain that which he cannot lose怎么理解

He is no fool who gives what he cannot keep to gain that which he cannot lose怎么理解
他不是个傻子,会给出他无法坚持赢得的东西,而这东西是他不能失去的.

who引导定语从句,what引导原因状语从句,that引导定语从句。